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The Ark is any of several ships featured in the Transformers franchise.
In The Transformers, the Ark was built by the Autobots to enable them to depart Cybertron due to the damage done to their planet by the Great War. They succeeded in launching it but were pursued by the Nemesis, whose crew boarded the Ark before it was caught in Earth's gravity well. The Ark subsequently crashed into the planet, knocking all of the inhabitants into emergency stasis. During the events of Beast Wars it was uncovered by the Maximals and Predacons, descendants of the Autobots and Decepticons, with Predacon Megatron attempting to kill Optimus Prime and alter history. This plan was thwarted by the Maximals, who established a base in the same volcanic mountain where the Ark had crash-landed and eventually managed to defeat Megatron.
A volcanic eruption in 1984 reawakened the slumbering Transformers, who resumed their conflict on Earth with the Ark serving as the Autobots' base. Sadly, it was later destroyed by the newly created Trypticon despite Autobot efforts to stop him.
The Ark appears in Transformers: Exodus, Transformers: Exiles, and Transformers: Retribution, where it is again the vessel used by Optimus Prime's crew to depart Cybertron. Having been built by Alpha Trion, it launches into space and-as in Generation 1-is pursued by the Nemesis. After passing through a Space Bridge, it arrives at Velocitron and is badly damaged by sabotage, though the Autobots are able to get it functioning again. They then travel to Junkion, where they engage in battle with the Decepticons and Star Seekers. Their path then takes them to Aquatron, where they are attacked by the Quintessons and the Ark is damaged, but manages to recover its crew from the planet before proceeding on its mission.